Biosemiotics and evolution: the natural foundations of meaning and Symbolism Elena Pagni Richard Theisen Simanke (Eds.) Introduction Paniel Reyes Cardenas Part I Life, meaning, and information 1. Exploring the philosophical background and scientific foundations of naturalist approaches to meaning and symbolism Richard Theisen Simanke Elena Pagni 2. Life sciences and the natural history of signs: can the origin of life processes coincide with the emergence of semiosis? Franco Giorgi 3. A proposal for a biosemiotic approach to digitalization: literacy as modelling competence Alin Olteanu 4. Threshold, meaning and life Arthur Araujo 5. How information gets its meaning Vinicius Romanini Part II Semiosis and evolution 6. Inclusive Fitness teleology and Darwinian explanatory pluralism: a theoretical sketch and an application to current controversies Philippe Huneman 7. The origins and evolution of design: a stage-based model Juan Mendoza-Collazos Jordan Zlatev Göran Sonesson 8. Biosemiotics and applied evolutionary epistemology: a comparison Marta Facoetti Nathalie Gontier 9. Extended synthesis and Jablonka and Lamb's four-dimensional view of evolution Jonathan Luís H. Ferreira Part III Physics, medicine, and bioenergetics 10. Physical intentionality: the phenomenological roots of biosemiotics Roberta Lanfredini 11. Cancer and cell death: a biosemiotic perspective Rogério Estevam Farias 12. Biosemiotics and bioenergetics: two perspectives compared Giulia Degl'Innocenti About the authors